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Ecology & Evolution

AB
Darwin's definition of evolutionDescent with modification over time
New synthesis definition of evolutionEvolution is change in allele frequencies of a population over generations
Two themes in The Origin of Species1) All organisms descended from a common ancestor with modifications. 2) The mechanism of modification over time is natural selection
Definition and three things needed for natural selection to occurNatural selection is a process in which individuals that have certain inherited traits tend to survive and reproduce at higher rates than other individuals because of those traits. Things needed: 1) Individuals must vary 2) Traits that vary must be heritable 3) Heritable traits must influence fitness
Predictions of Darwin's Theory: If species share common ancestry there should be traits shared by all species in a groupExample: homologous traits (traits that are similar due to common ancestor), such as the same bones existing in cat's paws/forelimbs and human's hands/arms
Predictions of Darwin's Theory: Natural selection should get rid of costly, unnecessary traitsExample: Emus are large, flightless birds that have vestigial wings that they do not use. Vestigial structures are remnants of features that surved a function on the organism's ancestors
Predictions of Darwin's Theory: If natural agents favor certain traits in certain environs, traits should be present in distantly related species who live in similar environsExample: Convergent evolution, the independent evolution of similar features in different lineages. N.American species (ie. flying squirrels) and Australian species (ie. sugar gliders) occupy similar environs and therefore look similar despite being very distantly related
Predictions of Darwin's Theory: We should see sequences of fossils globally that reflect sequential changes across earthExample: In the fossil record, the deeper the fossil the older the species. Closer to the surface, the species should have evolved. In shale dig sites in California, we can see this in how trilobites changed over time
Predictions of Darwin's Theory: Adult forms of species should be more divergent than embryonic formsExample: Early in development, we resemble distantly related species like cats (comparative embryology)
Predictions of Darwin's Theory: Humans should be able to direct evolution by selecting which adults produce offspringExample: Via artificial selection humans have been able to control the breeds of cats that arose; some for companionship and others for tasks like catching rats
Predictions of Darwin's Theory: Species should have limited distribution, only spreading via gradual migrationExample: There are no elephants in North America, but there are species of elephants in Asia and India. This shows that their spread has occurred but been limited
Predictions of Darwin's Theory: Because mutations arise regularly and accumulate over time, species related via narrow taxonomic groups (genus, family) should be more similar than those related via broad groups (genus, phylum)Example: Humans are more closely related to primates than we are to cats
Mendel's Law of SegregationWhen an individual produces gametes, the two alleles separate so that each gamete receives only one copy
Mendel's Law of Independent AssortmentEach pair of alleles segregates independently of each other pair of alleles during gamete formation
Mendalian inheritance assumes that: Genotype determines phenotypeException: Phenotypic plasticity. In hydrangeas, soil acidity influences the phenotype (flower color) due to phenotypic plasticity
Mendalian inheritance assumes that: Alleles are either dominant or recessiveException: Partial dominance and co-dominance. In partial dominance, both alleles are present, but one is blank. This can be seen in snapdragons, where red x white = pink. In Co-dominance, both alleles combine to make new colors. In parakeets, blue x yellow can create green
Mendalian inheritance assumes that: Each gene/locus affects only one traitException: Pleiotropy. In sickle cell disease, recessive allele at one locus affects both blood oxygen and severity of malarial infection. Alos, in siamese cats, the same protein produces abnormal coloration and crossed eyes
Mendalian inheritance assumes that: Each trait is only affected by one gene/locusException: Polygenic inheritance and epistasis. In human skin color, polygenic inheritance occurs because skin pigmentation is controlled by at least 3 separately inherited genes. In epistasis, the expression of a gene at one locus alters that of a gene at another. We see this in mice, because one allele can determine whether or not the mouse's fur has any color at all
Mendalian inheritance assumes that: All genes/loci sort independentlyException: Linkage. Linked genes are two or more genes on the same chromosome that tend to be inherited together
Mendalian inheritance assumes that: All traits in diploid organisms are affected by 2 copies of a geneException: Sex-linked traits (which refer to a single gene on a sex chromosome). Tortoiseshell cats can only be female and males can only be one color
Hardy-Wienburg equationp squared + 2pq + q squared = 1, where p represents the homozygous dominant alleles, 2pq represents the heterozygotes, and q represents the homozygous recessives
The five Hardy-Wienburg assumptions1) large populations 2) no mutations 3) no migration 4) no selection 5) mating is random, not selective
Violations of Hardy-Weinburg: Small populationA small population could cause low diversity, and is more susceptible to genetic drift and bottleneck effect
Bottleneck effectWhen few members of a species remain to mate, resulting in low genetic diversity. Example: Red wolves are facing a bottleneck effect in North Carolina, because the breeding program began with only 14 individuals
Genetic driftA process in which chance events can cause allele frequencies to fluctuate unpredictably from one generation to the next, especially in small populations
Founder effectWhen a few individuals become isolated from larger population, or found a new population whose gene pool differs from the source population. Example: Amish populations founded by a few members resulted in a disproportionately high number of rare recessive traits (polydactly combined with dwarfism )
Violations of Hardy-Weinburg: Mutations occurringWhen mutations occur they have the ability to create new alleles, which can throw off the H-W equilibrium
Violations of Hardy-Weinburg: Migration occurringMigration and gene flow involve individuals from other populations moving or mating with individuals from another population. This can throw off the allele frequencies. Example: Migration of new alleles in grass - some more tolerant to mineral concentrations in soil
Violations of Hardy-Weinburg: Selection occurringSelection violates the H-W assumption that mating is random. An assortative mating, individuals are picky about mates and select based on traits that contribute to fittness
Positive assortative mating"I prefer the cat that looks like me!!!" Results in fewer heterozygotes, more homozygotes
Negative assortative mating"I prefer different looking cats!!!" Results in more heterozygotes and fewer homozygotes
Evolution of resistance to pesticidesRed scale insects developed a resistance to hydrocyanide because when the insecticide was sprayed, the insects with the resistant allele survived to mate and pass it on to their offspring
Sexual selection: Male-male (intrasexual) competitionMales compete with other males to mate with the females. This is the most common form of sexual selection
Sexual selection: IntersexualFemales choose mate with the best display (ie. peacock's tail feathers). If a male has an impressive display, it's usually a good sign of his fitness
What is the only mechanism that leads directly to adaption?Selection
Darwin vs. MendelDarwin focues on population changes and patterns. Mendel focused on tracing traits across pedigrees
Mendel's four conclusions1) Inheritance is particulate, meaning that parents pass on "heritable factor" or gene 2) Variation is due to different alleles of a single gene 3) Each individual has 2 copies of a gene, one from each parent 4) One allele can be dominant and hide the other, recessive allele
TransmutationLamarck's (incorrect) theory that acquired traits could be inherited
Homozygous vs. heterozygousAn organism with a pair of identical alleles for a character is said to be homozygous (AA, aa). An organism that has two different alleles for a gene is said to be heterozygous (Aa)
Crossing overCrossing over occurs while replicated homologous chromosomes are paired during prophase of meiosis I and results in the end portions of two non-sister chromatids trading places, thus recombining linked genes
Catastrophism vs. uniformitarianismCatastrophism (Cuvier) is the principle that events in the past occurred suddenly and were caused by mechanisms different from those operating in the present. Uniformitarianism (Lyell) states that mechanisms of change are constant over time. Lyell's idea influenced Darwin's thinking



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